- The Paleontological Society
The Order Diaphanopterodea Handlirsch, 1919, is a small Paleozoic order of winged insects. For a long time, most of its species have been assigned to the order Megasecoptera. The position of the wings held backward along the abdomen at rest (Handlirsch, 1919, p. 575) is currently used to discriminate Diaphanopterodea from Megasecoptera (Carpenter, 1992). Although these Palaeoptera are frequently preserved with wings connected to the body, the wings resting position is not really suitable for ordinal diagnosis: isolated wings cannot be assigned, and taphonomic bias can render interpretations of wing disposal difficult. Here we redescribe Diaphanoptera munieri Brongniart, 1893, type-species of the Diaphanopteridae, and Diaphanoptera vetusta Brongniart, 1893. New diagnostic venation characters are established, some of which were already used for several others representatives of the order (Kukalová-Peck, 1974; Kukalová-Peck and Sinichenkova, 1992). The implications in the composition of the family and its relationships within the order are discussed.
In the following part we use the venation nomenclature of Kukalová-Peck (1991). Abbreviations: ScA, anterior Subcosta; ScP, posterior Subcosta; RA, anterior Radius; RP, posterior Radius; MA, anterior Media; MP, posterior Media; CuA, anterior Cubitus; CuP, posterior Cubitus; AA, anterior Anal.
Order Diaphanopterodea Handlirsch, 1906
Family Diaphanopteridae Handlirsch, 1906
Diaphanopteridae Handlirsch, 1906, p. 313; Carpenter, 1963, p. 241.
Diaphanopteritidae Handlirsch, 1919, p. 575; Rohdendorf, 1962, p. 69.
Diaphanoptera Brongniart, 1893.
ScA long parallel to anterior margin, distally vanishing in a pigmented (sclerotized?) apical area; area between anterior margin and ScA pigmented (sclerotized?); apex slightly pointed and slightly rejected posteriorly; ScP ending on RA just distal of midway of wing; few cross-veins between ScA and ScP, irregularly spaced, with one cross-vein just opposite end of ScP on RA; ScP and R basally posteriorly bowed; all cross-veins between RA and RP close to …