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Journal of Paleontology; January 2005; v. 79; no. 1; p. 110-124; DOI: 10.1666/0022-3360(2005)079<0110:CAITOP>2.0.CO;2
© 2005 Paleontological Society
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CHELICERATE ARTHROPODS, INCLUDING THE OLDEST PHALANGIOTARBID ARACHNID, FROM THE EARLY DEVONIAN (SIEGENIAN) OF THE RHENISH MASSIF, GERMANY

MARKUS POSCHMANN1, LYALL I. ANDERSON2 and JASON A. DUNLOP3

1 Department for the Protection of Cultural Monuments of Rhineland-Palatinate (Section for Geological Sites), Große Langgasse 29, D-55116 Mainz, Germany, <er-de.m.poschmann{at}t-online.de>,
2 Department of Geology and Zoology, National Museums of Scotland, Chambers Street, Edinburgh EH1 1JF, United Kingdom, <l.anderson{at}nms.ac.uk>,
3 Institut für Systematische Zoologie, Museum für Naturkunde der Humboldt-Universität zu Berlin, Invalidenstr. 43, D-10115 Berlin, Germany, <jason.dunlop{at}museum.hu-berlin.de>

A relatively diverse chelicerate fauna has been detected in Early Devonian, Siegenian strata of the Westerwald area, Rhineland-Palatinate, Germany. The arachnids, comprising trigonotarbids and the oldest phalangiotarbids, are described and figured here along with the chasmataspidids. To accomodate the phalangiotarbid a new genus and species in the family Architarbidae, Devonotarbus hombachensis, is raised. Devonotarbus n. gen. is characterized by an approximately straight posterior carapace margin, abbreviated and undivided anterior tergites, a large sixth tergite, and fused posterior tergites. The chasmataspidid closely resembles Diploaspis casteri from the Emsian assemblage of Alken an der Mosel, but is readily discernible as a new species, D. muelleri, by a strong tuberculation of its dorsal integument. With only fragmentary opisthosomal remains available, the trigonotarbids cannot be placed in known taxa with any certainty at this time.







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